Oviparous (Ref. 205). In captivity, spawning is preceded by vigorous driving by the male during the early hours of the day and eggs mostly sink to the bottom (Ref. 7020).
- Mills, D. and G. Vevers 1989 The Tetra encyclopedia of freshwater tropical aquarium fishes. Tetra Press, New Jersey. 208 p. (Ref. 7020)
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