Occurs inshore, on shelly ground, sand, muddy sand or mud. May enter freshwaters. Feeds on crustaceans (mysids, corophiid amphipods), small fish (gobies), bivalves, polychaetes, and chironomid larvae. Sexually mature at 1 to 2 years (Ref. 4696). Spawns in March to June with females spawning at least twice during this season. Adhesive eggs are deposited under or between stones, shells and aquatic plants and males guard the eggs until hatching (Ref. 59043).
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