Inhabits continental shelves from the littoral zone to about 300 m, where these may enter brackish water during early winter spawning migrations (Ref. 11954).
- Masuda, H., K. Amaoka, C. Araga, T. Uyeno and T. Yoshino 1984 The fishes of the Japanese Archipelago. Vol. 1. Tokai University Press, Tokyo, Japan. 437 p. (text). (Ref. 559) http://www.fishbase.org/references/FBRefSummary.php?id=559&speccode=7
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