Upstream spawning migration in summer to areas behind the peak of the flood. Spawning commences late in the afternoon from the surface to a depth of about 4 m, where there is a flow over a gravel, rock rubble substrate. Considerable spawning activity happens at the water surface. A single female circles slowly to the surface with her head deep and several males participate in courtship and finally in fertilization (Ref. 2906). Eggs are guarded and fanned by the male parent (Ref. 205).