Found in estuaries, lagoons and inshore waters over sand or mud, in seagrass or on algae (Ref. 4696). Occasionally recorded in freshwater, but there are no documented records of actual occurrence in European freshwaters (Ref. 59043). Feed on crustaceans (larger amphipods, isopods, shrimps, mysids, small crabs), bivalves, gastropods, polychaetes, chironomid larvae, sometimes small fish (Ref. 4696).