Found over hard bottoms. Mainly nocturnal, it feeds on small invertebrates. Oviparous (Ref. 205). Spawn in April (Marseille) or July (England); eggs are laid under mussel-shells or stones and guarded by the male (Ref. 5981).
- Wheeler, A. 1992 A list of the common and scientific names of fishes of the British Isles. J. Fish Biol. 41(1):1-37. (Ref. 5204) http://www.fishbase.org/references/FBRefSummary.php?id=5204&speccode=49
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