is the most wide-ranging species in the genus, with great variation in habitat, morphology, and chromosome number (G. Keator 1992). Two subspecies are recognized here, after G. Keator (1991, 1993b), but many other subspecies and even separate species have been recognized. For example, D. insulare has been recognized because it is endemic to the California Channel Islands and Guadalupe Island and is larger than mainland plants. Dichelostemma capitatum
ranges from northern Mexico through California, Oregon, Nevada, New Mexico, and possibly north to Idaho and Washington. Despite its great intraspecific variability, it is quite distinct from other species in the genus. It is the only species with six fertile stamens and epigeous germination, and it has unique corm, seed, and ovule characteristics. It also does not form hybrids with any other species, and it has even been placed in its own genus, Dipterostemon
(P. A. Rydberg 1912). Recent anatomical and molecular data support the idea that this species is not directly related to the other members of Dichelostemma
and perhaps is best treated as its own genus (R. Y. Berg 1996; J. C. Pires 2000).