Annual or perennial herbs. Leaves alternate. Capitula solitary, heterogamous; florets tubular; outer neuter; inner bisexual. Phyllaries many-seriate, imbricate, with a pectinate or spiny, apical appendage. Receptacle densely setose; setae twisted. Achenes ± laterally flattened, glabrous when mature. Pappus many-seriate, of persistent, unequal, barbellate, scale-like setae.
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