Dioecious trees. Leaves pinnately-veined, densely puberulous below on the reticulate venation. Stipules semi-amplexicaul, free. Inflorescences borne in leaf axils, unisexual, with both the male and female in the form of a spike. Male flowers with 4 tepals and stamens. Female flowers: tepals 4; stigmas 2, markedly unequal; fruiting perianth enlarged and fleshy, Fruit drupaceous.