Toco toucans are most commonly found in dry semi-open areas, which include regions such as woodland, savanna, plantations, and other regions that consist of scattered trees. In Brazil, toco toucans have been found in abundance in the "cerrado." Brazil's cerrado consists of savanna, semidecidious, and gallery forests surrounding river corridors. They are canopy frugivores that rely heavily on the availability of seasonal fruiting plants. Toco toucans therefore move from one habitat and region to the next in order to satisfy their dietary needs. This species is typically found at lowland elevations. However they have been sighted in elevations up to 1750 m around the Andes mountain range of South America.

Range elevation: 1750 (high) m.

Habitat Regions: tropical ; terrestrial

Terrestrial Biomes: savanna or grassland ; forest ; scrub forest

Other Habitat Features: riparian