Comments: This species has undergone considerable taxonomic revision in the past century and circumscription of subspecies remains a topic of considerable controversy. Workers in the field should be aware of the morphologic variation within the subspecies across the range of the species (i.e., approximately from the Sierra Nevada in the west to the plains of the Rocky Mountains in the east). Because rangeland managers and conservationists can often identify local morphologic and chemical races based on grazing or habitat preferences of wildlife and domestic animals, some impetus exists to further subdivide the subspecies at the varietal level. This treatment of the species complex remains conservative in light of the need for further study (Flora of North America Editorial Committee, 2006).
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