Description of parthenogenetic female: body elongate, spindle-shaped in dorsal view, in lateral view slightly arched dorsally and gradually tapering distally; shape fairly constant; transversal folds inconspicuous; colour pale olive-green to brown. Head indistinctly offset by neckfold, slightly tilted ventrally, broadly rounded anteriorly; chin weak; auricles absent; two very weak antero-lateral depressions; laterally from rostrum three pairs of tiny sensory papillae; dorsal antenna small, in anterior half of head. Lateral antennae small, the height of the bladder. Tail a weak lobe over cloacal opening. Foot rudimentary, short. Toes short, small, thin with short tubuli, conical in dorsal view; in lateral view dorsal margin slightly arched and ventral margin more or less straight. Caudal antenna distinct, dorsally near basis of toes. Brain large, obpyriform; eyespot large, dark red; retrocerebral sac hemispherical with few minute pigment granules. Oesophagus relatively long and narrow, annulation apparently absent. Stomach and intestine not separated by constriction. Gastric glands compound, composed of large, spherical ovate unstalked basal part attached to the stomach, and tubular anterior part connected proximally to mastax. Bladder small. Pedal glands fairly large, rounded ovate, with reservoir in foot. Vitellarium with eight nuclei; oviparous.
Trophi. Rami lyrate with distinct alulae and large openings proximally; latero-distally a small, narrow chamber with small opening dorso-caudally; inner margin of rami tips with scleropili. Fulcrum slightly less than ramus length, in lateral view almost parallel-sided, slightly tapering and rounded distally; no basal plate. Unci a thin striated, more or less rhomboid plate, composed of ± 15 tiny appressed teeth; ventral teeth largest; subuncus a tuft of loosely bound, relatively long scleropili. Manubria with characteristic ventral branch; ventral branch rod-shaped, slender, ~ ¾ shaft length and nearly parallel to it; head broad, plate-shaped, composed of three manubrial chambers lying next to each other; dorsal chamber with large opening at inner side of head; median and ventral chamber with small, opening at inner side of head near basis of shaft and ventral branch. Epipharynx a broad curved plate, and a pair of elongate^triangular platelets; epipharyngeal plate with lateral flaps delimited by a broad sinus proximally and a narrower incision distally; distally the central part of epipharyngeal plate bears two incurved branches, with smooth or slightly notched outer margin at the incurvation; the lateral flaps show a blunt apex distally; both incurved branches and apices are slightly more reinforced than the rest of the epipharyngeal plate, that shows the individual sclerite bodies it is built of; the exact position of the thin, elongate-triangular epipharyngeal platelets, composed of long sclerite bodies, could not be determined by light microscopy nor SEM.
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