Morphology
Body elongate, stout, fusiform in dorsal view, ventrally bent in lateral view; broadly oval in cross-section, higher than wide, greatest diameter near mid-length, fairly hyaline. Head and neck region narrower than trunk; head offset by dorsal transverse fold, occasionally with dorsal transverse fold anteriorly; neck offset from trunk by shallow, dorsal transverse fold. Trunk arched dorsally, more or less abruptly narrowing posteriorly; tail distinct, rounded posteriorly. Foot moderately short, c. 1/5 total length, two pseudosegments of equallellgth. Toes straight, more or less lanceolate in lateral view, abruptly ending in tubular points; in dorsal view with straight inner margins and curved outer margins; inner margins with short indentation prior to tubules, tubules laterally outcurved prior to their free end. Corona ventral, an anterior row of close-set cilia (reduced ·circumapical band ?) and at some distance (reduced apical field?) from the latter six tansverse, pre-oral bands of close-set cilia; lateral ciliary tufts absent. Dorsal antenna unpaired, short. Mouth near posterior edge of corona. Retrocerebral organ with sac and a pair of subcerebral glands. Brain egg-shaped, surrounded by large retrocerebral sac, two ducts, sac and ducts granular. Eyespots absent? No distinct constriction between stomach and intestine. Gastric glands medium large, globular, slightly compressed laterally, short-stalked. Pedal glands large, elongated, extending into trunk, with reservoir in toes. Vitellarium rounded, 8 nuclei. Subitaneous egg elongate oval in dorsal view, ventrally flattened in lateral view; short-stalked; smooth.
Trophi modified malleate. Rami triangular, inner margins, smooth, three to four short projections prior to tip; basal apophyses asymmetric: right apophyse small, 3toothed, left large, triangular, 7-toothed. Fulcrum short, in ventral view rod-shaped, slightly expanded posteriorly; in lateral view with expanded, ventrally recurved, hook-shaped posterior end. Left and right uncus with one principal and 5-6 subsidiary teeth, principal teeth with single accessory toothlet; principal teeth slightly clubbed, others linear, a supplementary comb of sharp, delicate teeth underneath tips of uncinal teeth. Manubria long, slender, strongly incurved posteriorly, head with short lateral cavities, anterior broadest . Two small, club-shaped epipharyngeal elements.
