The Woodland Brown is found on warm, open places in damp or mesic, deciduous or mixed woods with well-developed shrub and herbaceous layers. These habitats may be flooded in winter. The butterflies rarely visit flowers, preferring to feed on honeydew, moisture on buds and sap that runs from wounded trees. The males often settle on puddles on the ground, while the females tend to stay in the higher scrub. Females and caterpillars are restricted to a narrow zone under the tree and bush canopy along the edges of clearings where the host plant must be present. Eggs are laid on all species of grasses, mostly on false-bromes (Brachypodium spp.) but also on fescues (Festuca spp.), meadow-grasses (Poa spp.), small-reeds (Calamagrostis spp.) and on Sedges (Carex spp.). The half-grown caterpillar hibernates in a grass tussock, where later in the year it also pupates. The Woodland Brown has one generation a year. Habitats: broad-leaved deciduous forests (45%), mixed woodland (29%), alluvial and very wet forests and brush (8%), coniferous woodland (5%).
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