Inhabit rocky riffles of creeks and small to medium rivers (Ref. 5723, 10294); also found in small to medium upland streams (Ref. 10294). Adults feed on midge, blackfly, and caddisfly larvae, mayfly nymphs, isopods, and amphipods (Ref. 10294). Eggs are found clustered on underside of stone and guarded by males (Ref. 7043).
- Page, L.M. and B.M. Burr 1991 A field guide to freshwater fishes of North America north of Mexico. Houghton Mifflin Company, Boston. 432 p. (Ref. 5723) http://www.fishbase.org/references/FBRefSummary.php?id=5723&speccode=2590