Giant pandas scent-mark with urine and secretions from the anogenital region, using a variety of postures. Squatting and rubbing the perineum on a surface is common to both sexes. On a vertical surface, a leg lift may be used to permit anogenital contact and rubbing. The male often urinates and/or rubs the perineum in the leg-lift posture while the female generally only rubs the anogenital region. The female occasionally urinates in a 'handstand' position where both hind limbs are raised off the substrate; as a juvenile he would urinate and rub the anogenital region in a handstand. Both sexes sniffmarking sites extensively, and there is a noticeable build-up of secretions and discolouration at preferred locales.
- Kleiman, Devra G; Karesh, William B; Chu, Penn R. 1979. Behavioural changes associated with oestrus in the Giant panda Ailuropoda melanoleuca with comments on female proceptive behavior. International Zoo Yearbook Vol. 19, 217-223
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