Comments: Part of a complex formed by differing shell shapes and apparently different behaviors. Revision of this species group requires analysis of shell, antomical, and biochemical characters and must encompass all species of the genus Villosa. characters and must encompass all species of the genus Villosa. Due to much confusion surrounding the use of Unio nebulosus within the Villosa iris complex, Parmalee and Bogan (1998) chose to list all the described taxa from the Ohio, Tennessee, and Cumberland River systems as synonyms of Villosa iris and restrict the use of Villosa nebulosa to the species occurring in the headwaters of the Mobile Bay Basin.
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