Photograph from: Hamel-Leigue, A.C., S.K. Herzog, D.J. Mann, T.H. Larsen, B.D. Gill, W.D. Edmonds, & S. Spector. 2009. Distribution and natural history of the dung beetle tribe Phanaeini (Coleoptera: Scarabaeidae: Scarabaeinae) in Bolivia. Kempffiana, 5(2):43-95