With four occipital spines, laterals longer than medians. Posterior spines two, unequal and divergent; the right spine usually longer. Four occipital spines, the laterals longer than the medians. Right posterior spine longer than the left. Toes characteristically with claws. With four occipital spines; the laterals longer than the medians. Two unequal and divergent posterior spines, the right spine usually long.