The mangrove periwinkle, Littorina angulifera, is a common intertidal snail in mangrove forests of the southeast United States. The shell color of L. angulifera varies from bluish white, orange to dull yellow, reddish brown to grayish brown (Andrews 1994). The shell is comprised of 6 whorls, with the body whorl about half of the total height of the snail. Darker dashes on the ribs of the shell are often fused to form stripes on the body whorl. The early whorls around the base bear regularly-spaced vertical white spots below the channeled sutures.
- Kaplan, EH. 1988. A field guide to southeastern and Caribbean seashores: Cape Hatteras to the Gulf coast, Florida, and the Caribbean. Houghton Mifflin Co. Boston, MA. USA. 425 pp.
- Ruppert, EE & RD Barnes. Invertebrate zoology, 6th edition. Saunders College Publishing. Orlando, FL. USA. 1056 pp.
- Abbott, RT. 1974. American seashells: the marine Mollusca of the Atlantic and Pacific coasts of North America. Van Nostrand Reinhold Co. New York, NY. USA.
- Abbott, RT & PA Morris. 1995. A field guide to shells: Atlantic and Gulf coasts and the West Indies, 4th edition. Houghton Mifflin Co. Boston, MA. USA.
- Andrade, SCS & VN Solferini. 2006. The influence of size on the radula of Littoraria angulifera (Gastropoda: Littlorinidae). Malacologia. 49: 1-5.
- Andrews, J. 1994. A field guide to shells of the Florida coast. Gulf Publishing Co. Houston, Texas. USA. 182 pp.
- Chaves, AMR. 2002. Entre o seco e o molhado, do costão ao manguezal: distribuição de gastrópodes fa família Littorinidae em gradients vertical e horizontal no litoral do estado de São Paulo. Master's Thesis. Universidade Estadual de Campinas. Brazil.
- Gallagher, SB & GK Reid. 1979. Population dynamics and zonation in the periwinkle snail, Littorina angulifera, of the Tampa Bay, Florida region. Nautilus. 94: 162-178.
- Kohlmeyer, J & B Bebout. 1986. On the occurrence of marine fungi in the diet of Littorina angulifera and observations on the behavior of the periwinkle. Mar. Ecol. 7: 333-343.
- Merkt, RE & AM Ellison. 1998. Geographic and habitat-specific variation of Littoraria (Littorinopsis) angulifera (Lamarck, 1822). Malacologia. 40: 279-295.
- Tanaka, MO & RC Maia. 2006. Shell morphological variation of Littoraria angulifera among and within mangroves in NE Brazil. Hydrobiologia. 559: 193-202.
