"Umbrella to 1.2 m in diameter. Exumbrella: smooth, with mesoglea at center. Umbrellar margin: divided into 24 lappets more or less distinct; velar lappets: 8, broadly rounded, alternating with paired rhopaliar lappets: 16, lingulate. Rhopalia: 8, in deep niches between rhopaliar lappets. Tentacles: subumbrellar, in 8 adjacent groups, less than 10 per group, flattened, noticeably thick (3-5 mm or more in diameter), very long (greater than 5 mm), adjacent to periphery coronal muscle, arranged in a single linear row across proximal portion of velar lappets only. Coronal muscle: circular, folded, well-developed, divided by radial septa into 16 trapezoidal muscle fields; velar muscle fields alternate with somewhat narrower rhopaliar fields. Oral arms: 4, broad, curtainlike, pleated, about length of umbrella diameter or longer. Gonads: 4, large, saclike, everted, about 1/3 length of oral arms. Gastric pouches: 16, velar pouches slightly broader than rhopaliar pouches, terminating as blind-ending (rarely anastomosing) canals. Velar canals: dendritic, about equal in number at origin to tentacles, branching 3-4 times (mostly unilaterally) with branches becoming progressively shorter. Rhopaliar canals: single and broad at origin but narrowing toward umbrellar margin and branching bilaterally into short secondary and tertiary branches. Color: in preserved specimens, uniformly pink; in life, pink-violet."
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