Thallus: foliose, adnate to tightly adnate, 2-6 cm in diam., irregularly lobate; lobes: subirregular to irregular, flat, contiguous, 2-5 mm wide; apices: rotund; upper surface: yellow-green but darkening with age, dull in center but shiny at lobe apices, emaculate, smooth, becoming irregularly cracked and areolate in center, without soredia, isidia or pustulae; medulla: white; lower surface: shiny, pale ivory to light brown, darker at apices, moderately rhizinate; rhizines: simple, slender, concolorous with lower surface; Apothecia: sessile to subpedicellate, 2-5 mm wide; disc: dark brown, concave to flat, ultimately undulate-distorted; margin: smooth then crenate; ascospores: hyaline, simple, ellipsoid, 8-9 x 5-6 µm; Pycnidia: rare, immersed; conidia: bifusiform, 5-7 x c. 0.7 µm; Spot tests: upper cortex K-, C-, KC-, P-; medulla K-, C-, KC-, P-; Secondary metabolites: upper cortex with usnic acid (major); medulla with hypoprotocetraric acid (major), 4-O-demethylnotatic acid (submajor), conhypoprotocetraric acid (trace).; Substrate and ecology: on acidic rocks, often in open habitats; World distribution: North America and southern Africa; Sonoran distribution: infrequent at lower to intermediate elevations in Sonora.; Notes: Xanthoparmelia domokosioides is characterized by its tightly adnate, foliose thallus, its broad lobes with rotund apices, its pale lower surface, its lack of isidia and soredia and the production of hypoprotocetraric and 4-Odemethylnotatic acids in the medulla. Xanthoparmelia domokosioides resembles X. prodomokosii Hale, Elix & J. Johnst. as both have tightly adnate, foliose thalli with a pale lower surface and lack isidia and soredia, but these species differ in medullary chemistry and the width of the lobes. Whereas X. domokosioides has broad lobes (2-5 mm wide), larger spores (8-9 x 5-6 µm vs. 6-8 x 4-5 µm) and contains hypoprotocetraric and 4-O-demethylnotatic acids, X. prodomokosii has narrower lobes (1-2.5 mm wide), smaller spores and contains hypoprotocetraric acid and ±various scabrosin derivatives. The latter species appears to be confined to Australia whereas X. domokosioides is found in southern Africa and Mexico. Xanthoparmelia domokosii (Gyeln.) Hale is another species with identical chemistry to X. domokosioides, but differs in having a much small thallus and narrower lobes with a black lower surface.
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