Clarke (1980) described a subspecies, H. b. corpuscula, from whale stomach taken in the South Atlantic (type locality - 30°S,31°E). Voss et al., 1998, in an extensive study of the species, found that the characters Clarke used to separate the subspecies were insufficient. They suggest, however, that different populations of the species may exist based on differences in spermatophore structure. If this proves correct, they suggest that the Atlantic subtropical form could be identified to H. b. corpuscula.
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