Relationships of Caryophyllales to Other Clades of Eudicots
Relationships of Caryophyllales to other major clades of eudicots remain unclear. Cronquist (1981) and Takhtajan (1980, 1997), on the basis of floral characters, viewed Caryophyllidae as being derived from Ranunculales-type ancestors. However, phylogenetic analyses using many gene sequences place Caryophyllales firmly within the core eudicots. They have alternatively been considered to be close relatives of rosids, asterids, or Santalales and are best regarded at this time simply as one of the major clades of core eudicots (e.g., D. Soltis et al. 2000). The sister group of Caryophyllales may be Dilleniaceae (e.g., D. Soltis et al. 2000), although this relationship had not previously been suggested. Further investigation is needed to assess this potential relationship between Dilleniaceae and Caryophyllales.
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